| Index of Dragonflies Sabah | Index Dragonflies in Malaysia | Exoskeleton | LINKS | Simplified Chinese Version | Traditional Chinese Version | English Version |
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| Family Libellulidae | Red Dragonfly of Sabah | Frons of Dragonfly |
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The compound eyes of dragonflies More on comparing the size, colour, and shape of the eyes of dragonflies... |
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| A | B | C | D | E |
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| Eyes separated widely from other | Eyes meet partially along a seam | Eyes barely touch at a centre point | Eyes meet along a long seam | Eyes close but not quite touching |
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| Dragonflies are common in Malaysia and can be found all
around the globe in every continent except in Antartica.
Dragonfly are very fast flyers, can
be very colourful, harmless, timid and love water. We see them dipping
their tail into water, skimming on the surface, resting on twigs and
darting here and there and they are very difficult to capture.
T hey are harmless to humans but to other
insects, they are fearsome flying predator with the most advance
aviation system for aerial combat. If size doesn't matter, a dragonfly
can down the latest US jet fighter Raptor in seconds.
Dragonflies rely almost on their eyes to find a mate.
By middle of the day when temperatures are getting hot, males became
more active in their searches for females. |
| Associations between
dragonflies (Odonata) and their forest habitats in North Borneo:
implications for conservation. Albert George Orr ENS, Griffith University, Nathan Q 4111 ABSTRACT The Order Odonata (dragonflies and damselflies) includes nearly 6000 species worldwide. Owing to their charismatic appeal, ease of observation, and a life history which depends on both suitable freshwater and terrestrial habitats, they are increasingly carrying the flag for invertebrate conservation, both as indicators of environmental health, and as a group of special intrinsic value deserving priority protection in their own right. The greatest species richness of Odonata occurs in the Oriental and Neotropical regions, which boast respectively 1666 and 1636 described species. It is estimated that 25-30% of the total fauna is yet to be described in both regions. Within the Oriental region 23 of the 31 extant odonate families occur, 20 of which are found in Malaysia. At present about 360 species are recorded from Malaysia, 235 from West Malaysia and 275 from East Malaysia. Within Malaysia 80 species and 20 genera are known only from the peninsular, whereas 136 species and 10 genera are known only from North Borneo. However taken overall the Bornean fauna is much more distinct, with 46% species endemic to the island. It is believed that the greater part of the world tropical odonate fauna is dependent on forest habitats and unpolluted streams for its survival. The island of Borneo was originally almost completely covered by closed canopy tropical rainforest. Owing to an aseasonal, hot, per humid climate and high rainfall, forests were well supplied with streams and standing water. Consequently the rich, largely endemic odonate fauna must have evolved in association with these forests, and non-forest species, common today in disturbed land, must formerly have been rare opportunists in forest gaps or localized lacustrine species. It is estimated that at least 70 % of the fauna is presently confined to forest habitats and probably depends on forest for its survival. This study relates quantitatively odonate distribution to a mosaic of complex tropical rain forest formations in Brunei Darussalam. The tiny sultanate of Brunei still enjoys about 80% forest cover, representative of all the seven major formations found on the island and a great many of the 30+ sub-formations, and results from a nation-wide survey of odonates from most habitats are considered to be broadly applicable to the entire island of Borneo and many other parts of equatorial south-east Asia. Greatest odonate diversity, both Alpha and Beta, and greatest endemicity, is found in the primary lowland mixed dipterocarp forests, especially those growing in highly dissected landscapes such as occur at the Kuala Belalong Field Studies Centre, at the edges of the central uplands. High diversity and endemicity is also found in swamp forest, especially freshwater swamp, with certain endangered peat swamp formations also important. The highly vulnerable kerangas forest harbours fewer species, none uniquely, and the mangrove fauna is still more depauperate, with only a single wide-ranging specialist restricted to this habitat. Secondary dipterocarp forest is certainly less rich in odonates than primary forest, but lack of sites for parallel comparisons makes it difficult at present to state how serious this effect is. These results emphasize the importance of conserving a wide range of primary forest formations to achieve satisfactory odonate conservation, a strategy congruent with the conservation of charismatic land-based vertebrates and forest peoples. |
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Males and females can be distinguished as follows. Males have a bump containing the accessory genitalia under the second abdominal segment; females lack this. Females typically have a broader abdomen than males, and female damselflies and darners (Aeshnidae) have a prominent ovipositor under the last segment. Male dragonflies have 3 appendages at the abdomen tip, male damselflies 4; females of both have only 2 such appendages.
An important feature for identification in some families is the vulvar lamina under the 9th abdominal segment that supports the eggs about to be laid.
The females of all damselflies and some dragonflies (Aeshnidae and Petaluridae
families ) have a fully formed ovipositor, which is
a complicated structure containing paired valves and cutting blades, on the
underside of abdominal segments 8 and 9. The ovipositor is used to insert eggs
into plant tissue, mud, or other substrate.
Some species have a stylus, which is a thin, needle like projection, at the end of each of the two valves of the ovipositor.
Vulvar lamina with a conspicuous “V” incision
More on Ovipositor of dragonflies of Malaysia...
Vulvar lamina (subgenital plate/ (vulvar lamina).
Species without a true ovipositor have a more or less well developed vulvar lamina,
Vulvar lamina is a plate that extends rearward from segment 8 to cover part of the under surface of segment 9. This plate may be used to carry egg masses or aid in the dispersal of eggs.
In the spiketail family (Cordulegastridae), the vulvar lamina is highly modified to form a spikelike structure that inserts eggs, much as a true ovipositor might, into aquatic substrates. Some species of damselflies have a vulvar spine on the rear lower margin of segment 8 that projects over the genital opening at the base of segment 9.
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| Orthetrum testaceum | Gynacantha basiguttata (Selys, 1882) | Neurothemis terminata |
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| Matured male Orthetrum testaceum | Male Orthetrum testaceum |
| Body parts of a male dragonfly | Head parts of a dragonfly |
Tyriobapta torrida Kirby, 1889 |
A Survey on dragonflies of Sabah (I nformation of Dragonflies of North Borneo)
Sabah has large number of species of Malaysia dragonflies...
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Ictinogomphus decoratus melaenops (Selys, 1858) |
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Gynacantha basiguttata (Selys, 1882)This green dragonfly visit houses in the country side at night... |
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Anax panybeus Hagen, 1867Found in all the same situations as A. guttatus but is everywhere considerably rarer and more crepuscular in its habits... |
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Genus :
Neurothemis A visual comparison of the 3 most confused Red dragonflies in Sabah |
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| Neurothemis terminata terminata (Ris, 1911 ) |
Neurothemis fluctuans (Fabricius, 1793) |
Neurothemis ramburii ramburii (Kaup in Brauer, 1866) |
| Male : Bright Red | Male : Dark Red | Male : Bright Red |
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Bigger ( hw, 28-32 mm) |
Smaller (hw, 21-25 mm) |
Bigger ( hw, 28-32 mm) |
| Neurothemis terminata terminata (Ris, 1911 ) |
Neurothemis fluctuans (Fabricius, 1793) |
Neurothemis ramburii ramburii (Kaup in Brauer, 1866) |
| Neurothemis terminata copulat for about 30 seconds. A time too short for most photographers to snap a photo. |
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| hind wing 28-31 mm | Male hind wing 22-24mm Female hind wing 22-25mm |
hind wing 28-30 mm |
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| Above : Female Below : Female's Right Wing |
Above : Female
Neurothemis fluctuans Below : Female's Right Wing |
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Exuvia - Larval Exoskeleton of Dragonflies (empty cases of dragonfly nymphs) |
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Dragonflies and Damselflies living in Denmark. by Erland Refling Nielsen http://home1.stofanet.dk/erland_refling/danish_dragonflies.htm
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Dragonflies in West Malaysia West Malaysia hosts over 230 species of dragonflies and damselflies, which is roughly double the European total. Although few of these are endemic, many have restricted Sundaic distributions, occurring only in Malaysia, southern Thailand and Sumatra. Natural history tours included Fraser’s Hill and Taman Negara. Interesting area of peat swamp forest, a habitat seldom visited by tourists of any kind. A short distance from Kuala Lumpur is the old and charming British hill-station of Fraser’s Hill, small streams in and below the montane forest here are sure to yield a number of exciting species and birds are very easy to watch here. Small streams at mid-altitude are sure to hold the robust damselfly Devadatta argyoides, the golden-winged Euphaea ochracea and the widespread but glorious Aristocypha fenestrella. A key endemic to find here is the unmistakeable Calicnemia rectangulata, which is restricted to the mountain ranges of West Malaysia. Taman Negera is an area of incomparable humid rainforest. This is one of the oldest tracts of rainforest in the world and supports some quite fantastic beasts, including one of the largest dragonflies in the world, the monstrous Tetracanthagyna plagiata. Time will also be spent exploring some lowland peat swamp forest – a habitat which although depauperate in birds and mammals hosts an interesting and specialised Odonate fauna. Here is a good place to search for the rare and local damselfly Podolestes buwaldai. Inevitably during the trip we will spend some time around Kuala Lumpur, enjoying some of the finest food in the world (here the best of Malay, Chinese and Indian food is available) – a mouth-watering prospect for many. In Kuala Lumpur visit the ponds of the Botanical Gardens to look for the common and widespread species. |
Favourite habitat of dragonflies and damselflies :
| Trickle/Seepage | Forest Stream | Open Stream | River | Marsh | Pond/Lake | Forest Pond |
| Lower Stream | Mid Stream | Upper Stream |
| Shallow stream with sands and cobbles, water ran slowly and stagnant passing highly diversified habitats along river. Shrubs and grasses along the stream. Site totally exposed to the sunlight. | Provided varieties of substrates which consist of stones, rocky, cobbles and sandy. Water surface partially covered by canopies of trees and shrubs. Slow and fast flowing water. | Rich of substrates like stone and big rocky. Stream was too small with very fast flowing water, provided open site which is totally exposed to the sunlight, deep and very crystal clear water |
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Biography of Philip Powell Calvert (H.B. White) Pacific Northwest Dragonfly Migration Project (S. Valley) Toxicity tests on Stylurus amnicola (Great Lakes Fisheries Commission) |

List of common dragonflies in Sabah, Malaysia
| Dragonflies attract much
attention of naturalists for their vibrant colour. The showy striking
colour
of their body amazes many onlookers.
Odonates (dragonflies and damselflies) are insects which contains three Suborders: 1) Anisoptera (dragonflies), 2) Zygoptera (damselflies), 3) Anisozygoptera. The two former suborders of dragonflies and damselflies are widespread whereas Anisozygoptera is represented by one single species in Japan and one in the Himalayas. There are more than 5000 odonate species worldwide. In Malaysia more than 300 species have been found. Dragonflies are helping us in keeping down the numbers of mosquitoes and we tend to forget that mosquitoes form a large part of the diet of many predators including dragonflies and birds. Photography on dragonflies is now getting popular among the nature lovers. Morning is the best time for a beginner in photography dragonflies. Most dragonflies require that their flight muscles reach a certain minimum temperature before they are able to fly efficiently. Often insects are sluggish in the morning as they wait for the temperatures to rise enough to get their flight muscles warm. In a cool morning the dragonflies are much more approachable. And the dew condensed on the dragonflies wings often makes for a nice photo. In cool, dewcy mornings, one may find dragonflies covered with dew. They usually hide in a more sheltered spot, among weeds or cattails. They won't be going anywhere until morning sun fully warm them up. Dragonflies have favourite perches location. One can get as close as physically possible to the perching site and set up the camera. For macro close up work, a small F-stop is usually required (F16 or so). The dragonfly will leave during the setup, but will normally return within minutes landing on the same stem tip in the same physical position. Some dragonflies hunt into the late afternoon aided
by their excellent eye-sight. They perch on twigs, grass blades, or leaves at
the water’s edge, waiting to grab small insects that come within range. A dragonfly
usually perches with its body in a horizontal position. They
take an upright posture as if doing a handstand. It had
assumed this “obelisk posture” to cool itself by minimising the rays of the sun
ray to the body. Dragonfly are magnificent flying machines. It can fly at speeds of nearly 100kph. It can hover like a helicopter, fly vertically, stop or turn sharply in the midst of rapid flight, even fly backwards. It has the best eyesight among insects. |

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Feats and Facts |
Fossils of prehistoric dragonflies have been found with wingspans of 30 inches and body lengths up to 18 inches. |
Adults of Odonata can fly backwards or forwards and even hover like a helicopter. They can do these incredible aerial tricks because their two pairs of wings move independently of each other, in contrast to most other insects. |
The fastest recorded insect flight belongs to the Australian dragonfly, Austrophlbia costalis, which can reach 36 miles per hour over short distances. |
Some dragonflies can have between 10,000 and 50,000 individual eyes (commatidia) in each compound eye. |
The naiad shoots out its labium very fast to catch a prey, and the forward movement requires less than three one-hundredths of a second. |
The common green darner found in the United States, Anax junuios, migrates north into Canada each summer. In the fall, the offspring of these dragonflies will make their way back to the south. |
The largest damselfly, Megaloprepus coerulatus, has a wing spread of more than seven inches. This damselfly is found in Central and South America. |
A dragonfly nymph can use jet propulsion to move forward very quickly. By pulling water into the rectal chamber of its abdomen and then shooting it out. |
The word for dragonfly in Japanese is "tombo", but many years ago dragonflies were called "akitsu." Japan was once called Akitsushima, meaning "The Island of the Dragon-fly." |
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Odonates are insects with two large compound eyes, two pairs of delicate and membranous wings, and a long slender abdomen. Their life history occurs in two different stages: aquatic and aerial stages. The larval stage is spent in aquatic environment, and the adulthood is in terrestrial. Both the larvae and adults are good hunters. Prey may be stalked or ambushed. Dragonflies receive less attention due to their less economical significance compared to other insects such as butterflies. Yet, they are good indicator for aquatic ecosystem, and the larvae (nymphs) are proven to be good biocontrol of insect pests. |
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Adult dragonflies and damselflies catch and eat insects while flying. They eat anything small enough for them to handle, including flies, wasps, moths, and beetles. Adults often fly away from their nymphal home in the water to another area where they feed for several weeks before returning to the pond, lake, or stream to mate and lay eggs. Mating of dragonflies and damselflies is peculiar in that the mate holds the female behind the head while the female receives the male reproductive cells. Mating pairs will stay clasped for several minutes to several hours depand on species. They even fly while joined together. Dragonflies and damselflies undergo incomplete metamorphosis (egg-nymph-adult). The eggs are laid in water or in water plants and hatch into an aquatic nymph (naiad). The naiad stage lasts for a few weeks to almost five years. The nymphs must shed their skin, or molt, in order to grow in size, and they can molt 10 to 15 times before becoming mature. The mature nymph crawls out of the water onto the stem of an aquatic plant, where the adult emerges from the skin of the nymph, dries, and expands its wings, and flys away in search of food. Dragonflies are different from damselflies Dragonflies hold their wings out from the body when at rest. Damselflies rest with their wings held together over their backs. Dragonflies and damselflies wings are transperant with many veins. Most species in Sabah have beautiful color patterns. The coloration of the male dragonfly often differs from that of the female.
Dragonlfy's colours are due to pigmentation and will fade upon death. This is unlike other insects colours which reflective qualities of their cuticle, Their mouthparts include large mandibles for biting and chewing. They have small hairlike antennae. There are 5,000 species of dragonflies and damselflies. About 500 to 1500 species are to be described and named in the world.
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![]() A Guide to the Dragonflies of Borneo: Their Identification and Biology By: Orr A G The first guide to the dragonflies of Borneo Island. The most comprehensive coverage for any tropical region. 275 species (60%) occurring on the island are described and illustrated in photographs and 25 beautiful plates of 1/2-wing drawings. Chapters on biology, classification and ecology, as well as a complete checklist. 19.5 x 26.5 cm. Many more yet to be discovered, Borneo has one of the richest and most exciting dragonfly faunas in the world. More than 40% species found nowhere else, making Borneo the most distinctive sub-region of Sundaland. It is home to such spectacular species as Tetracanthagyna plagiata, the heaviest of all dragonflies, many beautiful picture-winged chlorocyphids and euphaeids, and high-altitude endemics such as Matronoides cyaneipennis restricted to Mount Kinabalu and nearby mountains.
The
text augments the illustrations and provides useful information on
biology.
Introductory chapters discuss structure and general biology, ecology and
conservation, faunistics and biogeography and collecting techniques and
photography. There is a complete and up to date checklist. Illustrated
keys to families of adults give the reader an understanding of the
structures used in classifying dragonflies and augment the usefulness of
the illustrations of entire insects. Main larval forms are shown. This
book will be useful not just in Borneo, but also in neighboring parts of
south-east Asia. |
A
Pocket Guide to Dragonflies of Peninsular Malaysia
AG Orr
127 pages, colour illus, map.
Dragonflies are among the most beautiful of insects. Peninsular Malaysia and Singapore are home to more than 230 species (twice the number found in Europe). They include some of the most exquisite forms found anywhere in the World. They occur wherever there is suitable fresh water habitat. Many species frequent the borders of garden ponds in our largest cities, perching on lily pads with glowing red bodies in striking contrast to their green platform. Others haunt swift clear streams in virgin rainforest, while yet others are confined to blackwater swamps. A few occur deep in the forest understorey, far from streams or pools, where they breed in the water accumulated in cavities in tree trunks. This book figures 98.7% of species known from Peninsular Malaysia and Singapore. For most, coloured drawings of the whole insect (omitting one pair of wings) are provided. Where necessary, coloured or monochrome drawings showing diagnostic features are also included. For some species, especially small Zygoptera (damselflies), only detailed structures are figured, as the general resemblance between close species is strong. A wide range of larval types is also figured. |
| Asahina S | A list of Odonata from Thailand (Part I-XXI) | Bosco Offset,599/182 Thailand,Bangkok:Soi Yoo Dee,Yannawa |
| Hemline M Pinratana A | Atlas of the Dragonflies of Thailand Distribution Maps by Provinces | Thailand Bangkok:Brothers of St.Gabriel |
| Fraser F C | The Fauna of British India Including Ceylon and Burma. Odonata | London:Taylor and Francis |
| Bishop J E | Limnology of Small Malayan River Gombak | Netherlands The Hague:Dr.W.JunkPublication House |
| Chovanec A | Dragonfly (Insecta: Odonata) as indicators of the ecological integrity of aquatic systems- A new assessment approach | Verh.Internat.Verein.Limnologica |
| Chovanec A Raab R | Dragonflies (Insecta, Odonata) and the ecological status of newly created wetlands: Example for long-term bioindication programmes | Limnologica |
| Corbet P S | A Biology of Dragonflies | London:Witherby |
| Corbet P S | Dragonflies: Behaviour and Ecology of Odonata | United Kingdom:Harley,Colchester |
| Dudgeon D | The influence of riparian vegetation on the functional organization of four Hong Kong stream communities | Hydrobiologia |
| Kovacs M | Biological Indicators in Environmental Protection | England:Ellis Horwood Limited |
| Krebs C J | Ecological Methodology | University of British Columbia:Harper Collins Publishers |
| Ludwig J A Reynolds J F | Statictical Ecology: A Primer on Methods and Computing | New York:John Wiley and Sons Inc |
| Milen M | Dragonflies (Odonata: Insecta) in the Bulgarian Wetlands: Current status, distribution and their importance as Bio-indicator[A]. The Asian Wetlands: Bringing Partnerships into Good Wetland Practices 2001 | Malaysia:Penerbit USM |
| Moore N W | Intra- and interspecific competition among dragonflies (Odonata): An account of observations and field experiments on population density in Dorset 1954-1960 | Journal Animal Ecology |
| Okudaira M Sugimura M Ishida S | Dragonflies of the Japanese Archipelago in color | Hokkaido:Hokkaido University Press |
| Osborn R Samways M J | Determinants of adult dragonfly assemblage pattern at new ponds in South Africa. | Odonatologica |
| Pinel-Alloul B Methot G Lapierre L | Macroinvertebrate Community as a Biological Indicator of Ecological and Toxicological Factors in Lake Saint Francois (Quebec) | Environmental Pollution |
| Pinratana A Kiauta B Hamalainen M | List of the Odonata of Thailand and annotated bibliography. | Thailand,Bangkok:Brothers of St.Gabriel |
| Raab R Chovanec A Wiener A K | Aspects of Habitat Selection by Adult Dragonflies at a Newly Created Pond in Vienna, Austria | Odonatologica |
| Schmidt E | Habitat inventerization, characterization and bioindication by a "representative spectrum of Odonata species (RSO)" | Odonatologica |
| Steytler N S Samways M J | Biotope selection by adult male dragonflies (Odonata) at an artificial lake created for insect conservation in South Africa | Biological Conservation |
| Strahler A N | Quantitative analysis of watershed geomorphology | Transaction American Geophysic Union |
| Sugimura M Ishida S Kijima K | Dragonflies of the Japanese Archiphelago in Color. Japan | Hokkaido:Hokkaido University Press |
| Suh A N Samways M J | Development of a dragonfly awareness trail in an African botanical garden | Biological Conservation |
| Wahizatul A | Some aspect of ecology of running water odonates and their potentials as indicator of environmental quality | Malaysia:Unpublished M.Sc.Thesis.Universiti Sains Malaysia |
| Watson J A L Theishinger G Abbey H M | The Australian Dragonflies, A Guide to the Identification, Distributions and Hhabitats of Australian Odonata. | Australia,CSIRO:Canberra and Melbourne |
| Widermuth H Spinner W | Visual cues in oviposition site selection by Somatochlora artica(Zetterstedt) (Anisoptera;Corduliidae) | Odonatologica |
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Odonata related sites in Britain and Ireland |
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Fast key to dragonfly species http://www.bio.georgiasouthern.edu 私の昆虫アルバム-トンボ編 : http://www5e.biglobe.ne.jp |
Nature / Environment:
Nature Society of Singapore
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Wild Singapore
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TeamSeaGrass
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Chek Jawa Mortality & Recruitment Project
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Raffles Museum of Biodiversity and Research
habitatnews.nus.edu.sg
RBZ Book Reviews
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Blue Water Volunteers
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Reef Ecology Study Team
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Wildfilms
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Ecology Asia
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Bird Ecology Study Group
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Sungei Buloh Wetland Reserve
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Urban Forest
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Leshon Loves Nature
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God's Wonderful Creation
wondercreation.blogspot.com
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mountan-n-sea2.blogspot.com
Where Discovery Begins
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Art in Wetlands
artinwetlands.wordpress.com
Gone Fishing - Casual Space for the Unhurried Soul
gonefishing.com.sg
Castle Beach
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Animal Concerns Research & Education Society (ACRES)
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Action for Singapore Dogs
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Nature books, journals, gifts, binoculars, etc.
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Friendly hostel stay in Singapore:
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Nature & Environment in S.E. Asia:
Climate Change Organisation
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Eyes On The Forest
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Tony Wu Underwater Photography
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Malaysian Nature Society
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Wild Asia
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Economy and Environment Program for South East Asia
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References Baker, R. L., and H. F. Clifford. 1982. Life cycle of an Enallagma boreale Selys population from the boreal forest of Alberta, Canada (Zygoptera: Coenagrionidae). Odonatologica 11(4):317-322. Calvert, P. P. 1902, in Calvert, P. P. 1901-1908. Odonata. In Biologia Centrali Americana: Insecta Neuroptera. R. H. Porter & Dulau & Co.: London. Dec 1902, p. 114. Calvert, P. P. 1919. Gundlach's work on the Odonata of Cuba: a critical study. Transactions of the American Entomological Society 45:335-396. Cannings, R. A. 1989. Enallagma basidens Calvert, a dragonfly new to Canada, with notes on the expansion of its range in North America (Zygoptera: Coenagrionidae). Notulae Odonatologicae 3(4):53-55. Charpentier, T. de. 1840. Libellulinae europaeae descriptae e depictae. Lipsiae, Leopold Voss. 180 pp. Donnelly, T. W. 1989. The status of Enallagma cyathigerum (Charp.) and E. vernale Gloyd in south-central New York (Zygoptera: Coenagrionidae). Odonatologica 18:373-378. Gloyd, L. K. 1943. Enallagma vernale, a new species of Odonata from Michigan. Occasional Papers of the Museum of Zoology, University of Michigan 479:1-8. Hagen, H. A. 1861. Synopsis of the neuroptera of North America, with a list of the South American species. Smithsonia Miscellaneous Collections 4:1-347. Ingham, B. R., and C. E. Jenner. 1976. Life histories of Enallagma hageni (Walsh) and E. aspersum (Hagen) (Zygoptera: Coenagrionidae). Odonatologica 5:331-345. Johannsson, O. E. 1978. Co-existence of larval Zygoptera (Odonata) common to the Norfolk Broads (U.K.). Oecologia 32:303-321. Kellicot, D. S. 1895. Catalogue of the Odonata of Ohio, Part 1. Journal of the Cincinnati Society of Natural History 17:195-216. Kormondy, E. J., and J. L. Gower. 1965. Life history variations in an association of Odonata. Ecology 46:882-886. Macan, T. T. 1964. The Odonata of a moorland fishpond. Int. Revue ges. Hydrobiol. 49:325-360. Morse, A. P. 1895. New North American Odonata. Psyche 7:207-211. O'Brien, M. F., and P. D. Pratt. 1999 (In press). Enallagma anna, a damselfly new to the Great Lakes region (Odonata: Coenagrionidae). The Great Lakes Entomologist 32(1). Pearlstone, P. S. M. 1973. The food of damselfly larvae in Marion Lake, British Columbia. Syesis 6:33-39. Say, T. 1839. Descriptions of new North American neuropterous insects and observations on some already described by (the late) Th. Say. Journal of the Academy of Natural Science of Philadelphia 8:9-46. Selys-Longchamps, E. de. 1875. Notes on Odonata from Newfoundland collected in 1874 by Mr. John Milne. Entomologists Monthly Magazine 11:241-243. Selys-Longchamps, E. de. 1876. Synopsis des agrionines, cinquième légion: Arion (suite). Le genre Agrion. Bulletin de l'Académie royale des Sciences de Belgique (2) 42:480-531. Walker, E. M. 1953. The Odonata of Canada and Alaska, Vol. 1. University of Toronto Press: Toronto, Ontario. xi + 292 pp. Walsh, B. D. 1862. List of the Pseudoneuroptera of Illinois contained in the cabinet of the writer, with descriptions of over forty new species, and notes on their structural affinities. Proceedings of the Entomological Society of Philadelphia 1862:361-402. Walsh, B. D. 1863. Observations on certain N. A. neuroptera by Hagen, M. D., of Konigsberg, Prussia; translated from the original French MS., and published by permission of the author, with notes and descriptions of about twenty new N. A. species of Pseudoneuroptera. Proceedings of the Entomological Society of Philadelphia 2:167-272. [note: see also Walsh, B. D. 1862 for original description of adults, which were named in Walsh 1863. Source: Westfall and May 1996 ). Westfall, M. J., Jr. and M. L. May. 1996. Damselflies of North America. Scientific Publishers: Gainesville, Florida. x + 650 pp. |
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NEW INDEX : Dragonfly May 10, 2008 01:21:01 PM |
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