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Family Libellulidae  | Red Dragonfly of Sabah  |  Frons of Dragonfly |


The compound eyes of dragonflies

More on comparing the size, colour, and shape of the eyes of dragonflies...

A B C D E
Eyes separated widely from other Eyes meet partially along a seam Eyes barely touch at a centre point Eyes meet along a long seam Eyes close but not quite touching

 

 

 
Dragonflies are common in Malaysia and can be found all around the globe in every continent except in Antartica.

Dragonfly are  very fast flyers, can be very colourful, harmless, timid and love water. We see them dipping their tail into water, skimming on the surface, resting on twigs and darting here and there and they are very difficult to capture. T hey are harmless to humans but to other insects, they are fearsome flying predator with the most advance aviation system for aerial combat. If size doesn't matter, a dragonfly can down the latest US jet fighter Raptor in seconds.

Speed : Can reach 60km per hour.
Range : Detected to cover a total distance of 137km per day.
Agility : Can instantly change direction while at high speed. Humans can't take that much G force.

 


Vision : Compound eyes of about 26,000 lenses with individual sensors capable of 360 deg vision. Can focus up to 20 feet. Focusing speed more than twice of humans.

Dragonflies rely almost on their eyes to find a mate.

 


Audio : Dragonflies are deaf. Therefore you do not need to whisper when you are near them.


Camouflage : C olours and patterns on dragonflies are NOT for camouflage. Dragonflies employ a kind of optical illusion called motion camouflage to trick their prey.
They can move in a certain pattern where the prey will see them as stationary, at a different location or very far but in fact they are actually already closing down fast on the prey.

 


There are 2 general types (Infra Order) of dragonflies :
1. The Zygoptera which is more commonly called Damselflies. When at rest their wings are folded back like most other flying insects. previously I didn't know damselflies are dragonflies as well.


2. The Anisoptera which is commonly known as the True Dragonflies. They are larger and when resting, their wings are opened to the sides.

All dragonflies have 2 pair of wings. And when in flight the two different pair move inverse way meaning from the front, we'll see they'll form an 'X'. That's where the Star Wars probably came up with the X wing fighter.

Where can we find them?

By middle of the day when temperatures are getting hot, males became more active in their searches for females.

Dragonflies spend most of their life in larvae form in a pond, lake or stream. Different species may prefer different type of habitat. Some prefer shady spots, some sunny, some still water, some flowing, some prefer clear water, some prefer muddy. They live only for a few weeks or months as an adult dragonfly compared to a few months or a few years as larvae. The larvae are predators too feeding on mosquito larvae, other insects, tadpoles and even small fish. As larvae or more specifically called nymph, they instead fall prey to larger fish and bigger nymphs. As adults dragonflies fall prey to fish, reptiles and amphibians usually during mating and laying eggs when they are most vulnerable. Some may get entangled in a spider web and become prey but unlike other insects, due to their speed and agility, they are not an easy catch for birds.

What does adult dragonflies feed on?
A dragonfly can eat up to 600 mosquitoes a day. They also eat other small insects like moth, butterflies, flies and sometimes each other when food are scarce. Dragonflies are not venomous and they kill and eat by ripping and tearing apart their prey usually head first. From their hunting habits we can separate dragonflies into Hawkers and Darters. Hawkers are often seen continuously flying in circles. They scan the area, attack, eat in flight and re attack. Darters can be seen resting on a twig or leaf and darts out every now and then to attack a prey flying by and then return to the twig.

 


Associations between dragonflies (Odonata) and their forest habitats in North Borneo: implications for conservation.

Albert George Orr
ENS, Griffith University, Nathan Q 4111

ABSTRACT

The Order Odonata (dragonflies and damselflies) includes nearly 6000 species worldwide. Owing to their charismatic appeal, ease of observation, and a life history which depends on both suitable freshwater and terrestrial habitats, they are increasingly carrying the flag for invertebrate conservation, both as indicators of environmental health, and as a group of special intrinsic value deserving priority protection in their own right.
The greatest species richness of Odonata occurs in the Oriental and Neotropical regions, which boast respectively 1666 and 1636 described species. It is estimated that 25-30% of the total fauna is yet to be described in both regions. Within the Oriental region 23 of the 31 extant odonate families occur, 20 of which are found in Malaysia. At present about 360 species are recorded from Malaysia, 235 from West Malaysia and 275 from East Malaysia. Within Malaysia 80 species and 20 genera are known only from the peninsular, whereas 136 species and 10 genera are known only from North Borneo. However taken overall the Bornean fauna is much more distinct, with 46% species endemic to the island.
It is believed that the greater part of the world tropical odonate fauna is dependent on forest habitats and unpolluted streams for its survival. The island of Borneo was originally almost completely covered by closed canopy tropical rainforest. Owing to an aseasonal, hot, per humid climate and high rainfall, forests were well supplied with streams and standing water. Consequently the rich, largely endemic odonate fauna must have evolved in association with these forests, and non-forest species, common today in disturbed land, must formerly have been rare opportunists in forest gaps or localized lacustrine species. It is estimated that at least 70 % of the fauna is presently confined to forest habitats and probably depends on forest for its survival. This study relates quantitatively odonate distribution to a mosaic of complex tropical rain forest formations in Brunei Darussalam. The tiny sultanate of Brunei still enjoys about 80% forest cover, representative of all the seven major formations found on the island and a great many of the 30+ sub-formations, and results from a nation-wide survey of odonates from most habitats are considered to be broadly applicable to the entire island of Borneo and many other parts of equatorial south-east Asia.
Greatest odonate diversity, both Alpha and Beta, and greatest endemicity, is found in the primary lowland mixed dipterocarp forests, especially those growing in highly dissected landscapes such as occur at the Kuala Belalong Field Studies Centre, at the edges of the central uplands. High diversity and endemicity is also found in swamp forest, especially freshwater swamp, with certain endangered peat swamp formations also important. The highly vulnerable kerangas forest harbours fewer species, none uniquely, and the mangrove fauna is still more depauperate, with only a single wide-ranging specialist restricted to this habitat. Secondary dipterocarp forest is certainly less rich in odonates than primary forest, but lack of sites for parallel comparisons makes it difficult at present to state how serious this effect is. These results emphasize the importance of conserving a wide range of primary forest formations to achieve satisfactory odonate conservation, a strategy congruent with the conservation of charismatic land-based vertebrates and forest peoples.

 

Deformed dragonflies can be a measure of the quality of water in lakes and rivers.

This is a female with pair deformed fore wing due to problems in emergence or water contamination.


Males and females can be distinguished as follows. Males have a bump containing the accessory genitalia under the second abdominal segment; females lack this. Females typically have a broader abdomen than males, and female damselflies and darners (Aeshnidae) have a prominent ovipositor under the last segment. Male dragonflies have 3 appendages at the abdomen tip, male damselflies 4; females of both have only 2 such appendages.

 

An important feature for identification in some families is the vulvar lamina under the 9th abdominal segment that supports the eggs about to be laid.


O vipositor


The females of all damselflies and some dragonflies (Aeshnidae and Petaluridae families ) have a fully formed ovipositor, which is a complicated structure containing paired valves and cutting blades, on the underside of abdominal segments 8 and 9. The ovipositor is used to insert eggs into plant tissue, mud, or other substrate.

Some species have a stylus, which is a thin, needle like projection, at the end of each of the two valves of the ovipositor.

Vulvar lamina with a conspicuous “V” incision

More on Ovipositor of dragonflies of Malaysia...


Vulvar lamina (subgenital plate/ (vulvar lamina).

Species without a true ovipositor  have a more or less well developed vulvar lamina,

Vulvar lamina is a plate that extends rearward from segment 8 to cover part of the under surface of segment 9. This plate may be used to carry egg masses or aid in the dispersal of eggs.

In the spiketail family (Cordulegastridae), the vulvar lamina is highly modified to form a spikelike structure that inserts eggs, much as a true ovipositor might, into aquatic substrates. Some species of damselflies have a vulvar spine on the rear lower margin of segment 8 that projects over the genital opening at the base of segment 9.




Orthetrum testaceum Gynacantha basiguttata (Selys, 1882) Neurothemis terminata

Matured male Orthetrum testaceum Male Orthetrum testaceum
Body parts of a male dragonfly Head parts of a dragonfly


 

Tyriobapta torrida Kirby, 1889






A Survey on dragonflies of Sabah  (I nformation of Dragonflies of North Borneo)

Sabah has large number of species of Malaysia  dragonflies...



Ictinogomphus decoratus melaenops (Selys, 1858)

Gynacantha basiguttata (Selys, 1882)

This green dragonfly visit houses in the country side at night...


Anax panybeus Hagen, 1867

Found in all the same situations as A. guttatus but is everywhere considerably rarer and more crepuscular in its habits...


Macrogomphus quadratus  large and widespread in Sabah. A female Macrogomphus quadratus has a body total length of 76mm


Genus : Neurothemis

A visual comparison of the 3 most confused Red dragonflies in Sabah

Neurothemis terminata terminata  (Ris, 1911 ) Neurothemis fluctuans
(Fabricius, 1793)
Neurothemis ramburii ramburii
(Kaup in Brauer, 1866)
Male : Bright Red Male : Dark Red Male : Bright Red
Bigger

( hw, 28-32 mm)

Smaller

(hw, 21-25 mm)

Bigger

( hw, 28-32 mm)

Neurothemis terminata terminata  (Ris, 1911 ) Neurothemis fluctuans
(Fabricius, 1793)
Neurothemis ramburii ramburii
(Kaup in Brauer, 1866)
Neurothemis terminata copulat for about 30 seconds.  A time too short for most photographers to snap a photo.
hind wing  28-31 mm Male  hind wing 22-24mm

Female  hind wing 22-25mm

hind wing 28-30 mm
Above : Female

Below : Female's Right Wing

Above : Female Neurothemis fluctuans

Below : Female's Right Wing


 

Useful Travel Maps  from http://www.sabahtourism.com


Sabah's West Coast Map  (149kb .Zip)



 

ODONATA IN THAILAND and CAMBODIA

All the pictures were photographed in pure natural conditions without any restriction to a dragonfly freedom http://pisum.bionet.nsc.ru

 


Exuvia - Larval Exoskeleton of Dragonflies

(empty cases of dragonfly nymphs)


Dragonflies and Damselflies living in Denmark.

by Erland Refling Nielsen

http://home1.stofanet.dk/erland_refling/danish_dragonflies.htm

 


Dragonflies in West Malaysia

West Malaysia hosts over 230 species of dragonflies and damselflies, which is roughly double the European total. Although few of these are endemic, many have restricted Sundaic distributions, occurring only in Malaysia, southern Thailand and Sumatra.

Natural history tours included Fraser’s Hill and Taman Negara.

Interesting area of peat swamp forest, a habitat seldom visited by tourists of any kind.

A short distance from Kuala Lumpur is the old and charming British hill-station of Fraser’s Hill, small streams in and below the montane forest here are sure to yield a number of exciting species and birds are very easy to watch here.

Small streams at mid-altitude are sure to hold the robust damselfly Devadatta argyoides, the golden-winged Euphaea ochracea and the widespread but glorious Aristocypha fenestrella.

A key endemic to find here is the unmistakeable Calicnemia rectangulata, which is restricted to the mountain ranges of West Malaysia.

Taman Negera is an area of incomparable humid rainforest. This is one of the oldest tracts of rainforest in the world and supports some quite fantastic beasts, including one of the largest dragonflies in the world, the monstrous Tetracanthagyna plagiata.

Time will also be spent exploring some lowland peat swamp forest – a habitat which although depauperate in birds and mammals hosts an interesting and specialised Odonate fauna.

Here is a good place to  search for the rare and local damselfly Podolestes buwaldai.

Inevitably during the trip we will spend some time around Kuala Lumpur, enjoying some of the finest food in the world (here the best of Malay, Chinese and Indian food is available) – a mouth-watering prospect for many.

In Kuala Lumpur  visit the ponds of the Botanical Gardens to look for the common and widespread species.


Favourite habitat of dragonflies and damselflies :

             
Trickle/Seepage Forest Stream Open Stream River Marsh Pond/Lake Forest Pond

     
Lower Stream   Mid Stream Upper Stream
Shallow stream with sands and cobbles, water ran slowly and stagnant passing highly diversified habitats along river. Shrubs and grasses along the stream. Site totally exposed to the sunlight. Provided varieties of substrates which consist of stones, rocky, cobbles and sandy. Water surface partially covered by canopies of trees and shrubs. Slow and fast flowing water. Rich of substrates like stone and big rocky. Stream was too small with very fast flowing water, provided open site which is totally exposed to the sunlight, deep and very crystal clear water

Biography of Philip Powell Calvert (H.B. White)

Pacific Northwest Dragonfly Migration Project (S. Valley)

Toxicity tests on Stylurus amnicola (Great Lakes Fisheries Commission)

Dragonflies of Nova Scotia (P. Brunelle)

Regional Lists of Ontario Odonata (P. Pratt)


Neurothemis fluctuans Rhyothemis triangularis (Kirby, 1889)  三角蜻蜓 Trithemis festiva (Rambur, 1842) 庆褐蜻 (樂仙蜻蜓)

List of common dragonflies in Sabah, Malaysia

Dragonflies attract much attention of naturalists for their vibrant colour.  The showy striking colour of their body amazes many onlookers.

Odonates (dragonflies and damselflies)  are insects  which contains three Suborders: 1) Anisoptera (dragonflies), 2) Zygoptera (damselflies), 3) Anisozygoptera.

The two former suborders of dragonflies and damselflies are widespread whereas Anisozygoptera is represented by one single species in Japan and one in the Himalayas. 

There are more than 5000  odonate species worldwide. In Malaysia more than 300 species have been found. Dragonflies  are helping  us in  keeping down the numbers of mosquitoes and we tend to forget that mosquitoes form a large part of the diet of many predators including dragonflies and birds.

Photography on dragonflies is now getting popular among the nature lovers. Morning is the best time for a beginner in photography dragonflies. Most dragonflies  require that their flight muscles reach a certain minimum temperature before they are able to fly efficiently. Often insects are sluggish in the morning as they wait for the temperatures to rise enough to get their flight muscles warm. In a cool morning the dragonflies are much more approachable. And the dew condensed on the dragonflies wings often makes for a nice photo.

In  cool, dewcy mornings, one  may find dragonflies covered with dew. They usually hide in a more sheltered spot, among weeds or cattails. They won't be going anywhere until morning sun fully warm them up.

Dragonflies  have favourite perches location. One can get as close as physically possible to the perching site and set up the camera. For macro close up work, a small F-stop is usually required (F16 or so). The dragonfly will leave during the setup, but will normally return within minutes  landing on the same stem tip in the same physical position.

Some dragonflies hunt into the late afternoon aided by their excellent eye-sight. They perch on twigs, grass blades, or leaves at the water’s edge, waiting to grab small insects that come within range. A dragonfly usually perches with its body in a horizontal position. They  take  an upright posture as if  doing a handstand. It had assumed this “obelisk posture” to cool itself by minimising the rays of the sun ray to the body.
Dragonflies are usually found near ponds, lakes, rivers, or even puddles of water that are quite permanent. When they are not flying, you can find them perching on leaves or twigs at the water’s edge.

Dragonfly are magnificent flying machines. It can fly at speeds of nearly 100kph. It can hover like a helicopter, fly vertically, stop or turn sharply in the midst of rapid flight, even fly backwards. It has the best eyesight among insects.




Brachythemis contaminata (Fabricius,1793) Trithemis festiva (Rambur, 1842)

 


 

Feats and  Facts


Fossils of prehistoric dragonflies have been found with wingspans of 30 inches and body lengths up to 18 inches.

Adults of Odonata can fly backwards or forwards and even hover like a helicopter. They can do these incredible aerial tricks because their two pairs of wings move independently of each other, in contrast to most other insects.

The fastest recorded insect flight belongs to the Australian dragonfly, Austrophlbia costalis, which can reach 36 miles per hour over short distances.

Some dragonflies can have between 10,000 and 50,000 individual eyes (commatidia) in each compound eye.

The naiad shoots out its labium very fast to catch a prey, and the forward movement requires less than three one-hundredths of a second.

The common green darner found in the United States, Anax junuios, migrates north into Canada each summer. In the fall, the offspring of these dragonflies will make their way back to the south.

The largest damselfly, Megaloprepus coerulatus, has a wing spread of more than seven inches. This damselfly is found in Central and South America.

A dragonfly nymph can use jet propulsion to move forward very quickly. By pulling water into the rectal chamber of its abdomen and then shooting it out.

The word for dragonfly in Japanese is "tombo", but many years ago dragonflies were called "akitsu." Japan was once called Akitsushima, meaning "The Island of the Dragon-fly."

Odonates are insects with two large compound eyes, two pairs of delicate and membranous wings, and a long slender abdomen. Their life history occurs in two different stages: aquatic and aerial stages. The larval stage is spent in aquatic environment, and the adulthood is in terrestrial.

Both the larvae and adults are good hunters. Prey may be stalked or ambushed.

Dragonflies receive less attention due to their less economical significance compared to other insects such as butterflies.

Yet, they are good  indicator for aquatic ecosystem, and the larvae (nymphs) are proven to be good biocontrol of insect pests.


Adult dragonflies and damselflies catch and eat insects while flying. They eat anything small enough for them to handle, including flies, wasps, moths, and beetles. Adults often fly  away from their nymphal home in the water to another area where they feed for several weeks before returning to the pond, lake, or stream to mate and lay eggs.


Mating of dragonflies and damselflies is peculiar in that the mate holds the female behind the head while the female receives the male reproductive cells.

Mating pairs will stay clasped for several minutes to several hours depand on species. They even fly while joined together.


Dragonflies and damselflies undergo incomplete metamorphosis (egg-nymph-adult). The eggs are laid in water or in water plants and hatch into an aquatic nymph (naiad). The naiad stage lasts for a few weeks to almost five years. The nymphs must shed their skin, or molt, in order to grow in size, and they can molt 10 to 15 times before becoming mature. The mature nymph crawls out of the water onto the stem of an aquatic plant, where the adult emerges from the skin of the nymph, dries, and expands its wings, and flys away in search of food.


Dragonflies are different from damselflies

 Dragonflies hold their wings out from the body when at rest.

 Damselflies rest with their wings held together over their backs.


Dragonflies and damselflies wings are transperant with many veins. Most species in Sabah have beautiful color patterns. The coloration of the male dragonfly often differs from that of the female.

 

Dragonlfy's colours are due to pigmentation and will fade upon death. This is unlike other insects colours which reflective qualities of their cuticle,


Their mouthparts include large mandibles for biting and chewing. They have small hairlike antennae.


There are 5,000 species of dragonflies and damselflies. About 500 to 1500 species are to be described and named in the world.

 


Dragonflies and damselflies have large compound eyes that can see in all directions. When the compound eye is magnified several hundred times, each individual facet (ommatidium) is shown to be hexagonal in shape.

Photo Left : Compound eyes of a male  Crocothemis servilia servilia (Drury, 1770)


A Guide to the Dragonflies of Borneo: Their Identification and Biology By: Orr A G

The first guide to the dragonflies of Borneo Island. The most comprehensive coverage for any tropical region. 275 species (60%) occurring on the island are described and illustrated in photographs and 25 beautiful plates of 1/2-wing drawings. Chapters on biology, classification and ecology, as well as a complete checklist. 19.5 x 26.5 cm.

Many more yet to be discovered, Borneo has one of the richest and most exciting dragonfly faunas in the world. More than 40% species found nowhere else, making Borneo the most distinctive sub-region of Sundaland. It is home to such spectacular species as Tetracanthagyna plagiata, the heaviest of all dragonflies, many beautiful picture-winged chlorocyphids and euphaeids, and high-altitude endemics such as Matronoides cyaneipennis restricted to Mount Kinabalu and nearby mountains.

 
The first guide to Borneo’s dragonflies, is also the most comprehensively illustrated account of any large tropical dragonfly fauna yet published. Species are figured by natural photographs and half-wing drawings. About 60% of known species are shown, including almost all the distinctive and common species likely to be encountered by a casual visitor. Particular attention is given to the identification of the common but difficult medium-sized red dragonflies of which there are several.

The text augments the illustrations and provides useful information on biology. Introductory chapters discuss structure and general biology, ecology and conservation, faunistics and biogeography and collecting techniques and photography. There is a complete and up to date checklist. Illustrated keys to families of adults give the reader an understanding of the structures used in classifying dragonflies and augment the usefulness of the illustrations of entire insects. Main larval forms are shown. This book will be useful not just in Borneo, but also in neighboring parts of south-east Asia.
 

A Pocket Guide to Dragonflies of Peninsular MalaysiaA Pocket Guide to Dragonflies of Peninsular Malaysia AG Orr

127 pages, colour illus, map.
Natural History Publications
 

 

Dragonflies are among the most beautiful of insects. Peninsular Malaysia and Singapore are home to more than 230 species (twice the number found in Europe). They include some of the most exquisite forms found anywhere in the World. They occur wherever there is suitable fresh water habitat. Many species frequent the borders of garden ponds in our largest cities, perching on lily pads with glowing red bodies in striking contrast to their green platform. Others haunt swift clear streams in virgin rainforest, while yet others are confined to blackwater swamps. A few occur deep in the forest understorey, far from streams or pools, where they breed in the water accumulated in cavities in tree trunks. This book figures 98.7% of species known from Peninsular Malaysia and Singapore. For most, coloured drawings of the whole insect (omitting one pair of wings) are provided. Where necessary, coloured or monochrome drawings showing diagnostic features are also included. For some species, especially small Zygoptera (damselflies), only detailed structures are figured, as the general resemblance between close species is strong. A wide range of larval types is also figured.

Asahina S A list of Odonata from Thailand (Part I-XXI) Bosco Offset,599/182 Thailand,Bangkok:Soi Yoo Dee,Yannawa
Hemline M Pinratana A Atlas of the Dragonflies of Thailand Distribution Maps by Provinces Thailand Bangkok:Brothers of St.Gabriel
Fraser F C The Fauna of British India Including Ceylon and Burma. Odonata London:Taylor and Francis

Bishop J E Limnology of Small Malayan River Gombak Netherlands The Hague:Dr.W.JunkPublication House
Chovanec A Dragonfly (Insecta: Odonata) as indicators of the ecological integrity of aquatic systems- A new assessment approach Verh.Internat.Verein.Limnologica
Chovanec A Raab R Dragonflies (Insecta, Odonata) and the ecological status of newly created wetlands: Example for long-term bioindication programmes Limnologica
Corbet P S A Biology of Dragonflies London:Witherby
Corbet P S Dragonflies: Behaviour and Ecology of Odonata United Kingdom:Harley,Colchester
Dudgeon D The influence of riparian vegetation on the functional organization of four Hong Kong stream communities Hydrobiologia
Kovacs M Biological Indicators in Environmental Protection England:Ellis Horwood Limited
Krebs C J Ecological Methodology University of British Columbia:Harper Collins Publishers
Ludwig J A Reynolds J F Statictical Ecology: A Primer on Methods and Computing New York:John Wiley and Sons Inc
Milen M Dragonflies (Odonata: Insecta) in the Bulgarian Wetlands: Current status, distribution and their importance as Bio-indicator[A]. The Asian Wetlands: Bringing Partnerships into Good Wetland Practices 2001 Malaysia:Penerbit USM
Moore N W Intra- and interspecific competition among dragonflies (Odonata): An account of observations and field experiments on population density in Dorset 1954-1960 Journal Animal Ecology
Okudaira M Sugimura M Ishida S Dragonflies of the Japanese Archipelago in color Hokkaido:Hokkaido University Press
Osborn R Samways M J Determinants of adult dragonfly assemblage pattern at new ponds in South Africa. Odonatologica
Pinel-Alloul B Methot G Lapierre L Macroinvertebrate Community as a Biological Indicator of Ecological and Toxicological Factors in Lake Saint Francois (Quebec) Environmental Pollution
Pinratana A Kiauta B Hamalainen M List of the Odonata of Thailand and annotated bibliography. Thailand,Bangkok:Brothers of St.Gabriel
Raab R Chovanec A Wiener A K Aspects of Habitat Selection by Adult Dragonflies at a Newly Created Pond in Vienna, Austria Odonatologica
Schmidt E Habitat inventerization, characterization and bioindication by a "representative spectrum of Odonata species (RSO)" Odonatologica
Steytler N S Samways M J Biotope selection by adult male dragonflies (Odonata) at an artificial lake created for insect conservation in South Africa Biological Conservation
Strahler A N Quantitative analysis of watershed geomorphology Transaction American Geophysic Union
Sugimura M Ishida S Kijima K Dragonflies of the Japanese Archiphelago in Color. Japan Hokkaido:Hokkaido University Press
Suh A N Samways M J Development of a dragonfly awareness trail in an African botanical garden Biological Conservation
Wahizatul A  Some aspect of ecology of running water odonates and their potentials as indicator of environmental quality Malaysia:Unpublished M.Sc.Thesis.Universiti Sains Malaysia
Watson J A L Theishinger G Abbey H M The Australian Dragonflies, A Guide to the Identification, Distributions and Hhabitats of Australian Odonata. Australia,CSIRO:Canberra and Melbourne
Widermuth H Spinner W Visual cues in oviposition site selection by Somatochlora artica(Zetterstedt) (Anisoptera;Corduliidae) Odonatologica

Odonata related sites in Britain and Ireland

The Cambridgeshire Dragonfly Group
Cheshire Dragonflies & Damselflies
Dragonflies of the Hampshire and Surrey Borders
Hertfordshire Dragonfly Group
Lincolnshire Wildlife Trust
The Dragonfly Project
Shetland Wildlife
"Backyard Dragons"
Dragonflies and Damselflies in the UK - particularly in North Buckinghamshire
Bedfordshire Odonata
Dragonflies and Damselflies of Wimbledon and Putney Commons
Wildlife in the Knaresborough area (N. Yorks)
Warwickshire Dragonfly Group
Dragonfly Ireland project
Stover Country Park, Devon
Derbyshire Dragonflies
Northamptonshire Dragonflies
Leicestershire & Rutland Dragonfly Group
Staffordshire Ecological Record - A Survey of the Dragonflies of Lichfield
Yorkshire branch of the British Dragonfly Society
Dragonflies of the East Riding of Yorkshire
Dragonflies and Damselflies of Rochdale

Dragonflies and Habitat in Switzerland (B. Oertli )
Dragonflies and Damselflies of India (F. K. Kakkassery
R.J. Tillyard 1917 The Biology of Dragonflies

Stylurus plagiatus Adult (B. Barber)
Stylurus plagiatus Nymph (B. Barber)
Entomological Equipment, Nets, Envelopes, etc. (Hillside Books)
Odonata Publications for Sale (Hillside Books)
Catching and Preserving Dragonflies FAQ (Terry Morse)
Cameroon Dragonfly Project (David Chelmick)
Odonata Photos by William Zittrich
Atropos
Joshua Stuart Rose's Home Page
Advice for Teachers, Class Project, Monitoring Migratory Dragonflies (T. Morse)
Guidelines for Monitoring Migratory Dragonflies (T. Morse)

Fast key to dragonfly species  http://www.bio.georgiasouthern.edu

私の昆虫アルバム-トンボ編  : http://www5e.biglobe.ne.jp


Nature / Environment:
Nature Society of Singapore
www.nss.org.sg

Wild Singapore
www.wildsingapore.com
TeamSeaGrass
teamseagrass.blogspot.com
Chek Jawa Mortality & Recruitment Project
cjproject.blogspot.com
Raffles Museum of Biodiversity and Research
habitatnews.nus.edu.sg
RBZ Book Reviews
rbz-bookreviews.blogspot.com
Blue Water Volunteers
www.bluewatervolunteers.org
Reef Ecology Study Team
coralreef.nus.edu.sg
Pulau Hantu Blog
www.pulauhantu.org
Wildlife Singapore
www.wildsingapore.per.sg
Wildfilms
wildfilms.blogspot.com
Naked Hermit Crabs
nakedhermitcrabs.blogspot.com
Ecology Asia
www.ecologyasia.com
Singapore Snakes
singaporesnakes.blogspot.com
Bird Ecology Study Group
besgroup.blogspot.com
Eco Singapore
www.eco-singapore.org
Sungei Buloh Wetland Reserve
www.sbwr.org.sg
B-Pals (On-line Community for Butterfly Enthusaists)
www.b-pals.com

Nature Photography Society
www.naturephotosociety.org.sg
Nature Spies
naturespies.blogspot.com
Simply Green
www.simplygreen.com.sg
 

Singapore Adventurers' Club
www.sac.org
Earthlink - NTU
www.clubs.ntu.edu.sg/earthlink
Environmental News Archive
www.environmentalnews.blogspot.com

Kindred Spirits, Nature's Friends:

GreenTheme
greentheme.blogspot.com
The Annotated Budak
budak.blogs.com
Midnight Monkey Monitor
leafmonkey.blogspot.com
Colourful Clouds
colorclouds.blogspot.com
Lost in the Jungle
tuskerman.blogspot.com
The Tide Chaser
tidechaser.blogspot.com
Manta Blog
mantamola.blogspot.com
Urban Forest
uforest.blogspot.com
Leshon Loves Nature
leshonlovesnature.blogspot.com
God's Wonderful Creation
wondercreation.blogspot.com
Mountain & Sea
mountan-n-sea2.blogspot.com
Where Discovery Begins
wherediscoverybegins.blogspot.com
Art in Wetlands
artinwetlands.wordpress.com
Gone Fishing - Casual Space for the Unhurried Soul
gonefishing.com.sg
Castle Beach
www.castlebeach.org

Rights and welfare of animals:
Society for the Prevention of Cruelty to Animals (SPCA)
www.spca.org.sg
Animal Concerns Research & Education Society (ACRES)
www.acres.org.sg
Cat Welfare Society
www.catwelfare.org
Action for Singapore Dogs
www.asdsingapore.com
Animal Lovers League
www.animalloversleague.com
Animal Watch
www.animalwatch.org
House Rabbit Society Singapore
www.hrss.net
SOS Animals
www.sosanimals.com

Nature books, journals, gifts, binoculars, etc.
Nature's Niche
www.naturesniche.com

Natural products / lifestyle for a good cause:
Green Circle Eco-Farm
www.greencircle.com.sg
Kampung Senang Charity & Education Foundation
www.kg-senang.org.sg
Working for disadvantaged women in Nepal
www.simplynatural.com.sg
Vegetarian Society (Singapore)
www.vegetarian-society.org

History and Sights of Singapore:
Yesterday Singapore
www.yesterday.sg
Virtual tours of People, Places, Festivals and Nature of Singapore
www.singaporesights.com
Singapore Photopoetry
singaporephotopoetry.blogspot.com

Friendly hostel stay in Singapore:
HostelOne66
www.hostelOne66.com

Singapore College of Traditional Chinese Medicine:
www.singaporetcm.com

Nature & Environment in S.E. Asia:
Climate Change Organisation
www.cco-asia.org
Eyes On The Forest
www.eyesontheforest.or.id
Tony Wu Underwater Photography
www.tony-wu.com
Malaysian Nature Society
www.mns.org.my
Wildlife Asia
www.wildlifeasia.org
Wild Asia
www.wildasia.net
Avian Watch Asia
www.avianwatchasia.org
Sumatran Orang Utan Conservation Programme
www.sumatranorangutan.com
TRAFFIC Southeast Asia
www.traffic.org
Mangrove Action Project
www.mangroveactionproject.org
Economy and Environment Program for South East Asia
www.eepsea.org


References

Baker, R. L., and H. F. Clifford. 1982. Life cycle of an Enallagma boreale Selys population from the boreal forest of Alberta, Canada (Zygoptera: Coenagrionidae). Odonatologica 11(4):317-322.

Calvert, P. P. 1902, in Calvert, P. P. 1901-1908. Odonata. In Biologia Centrali Americana: Insecta Neuroptera. R. H. Porter & Dulau & Co.: London. Dec 1902, p. 114.

Calvert, P. P. 1919. Gundlach's work on the Odonata of Cuba: a critical study. Transactions of the American Entomological Society 45:335-396.

Cannings, R. A. 1989. Enallagma basidens Calvert, a dragonfly new to Canada, with notes on the expansion of its range in North America (Zygoptera: Coenagrionidae). Notulae Odonatologicae 3(4):53-55.

Charpentier, T. de. 1840. Libellulinae europaeae descriptae e depictae. Lipsiae, Leopold Voss. 180 pp.

Donnelly, T. W. 1989. The status of Enallagma cyathigerum (Charp.) and E. vernale Gloyd in south-central New York (Zygoptera: Coenagrionidae). Odonatologica 18:373-378.

Gloyd, L. K. 1943. Enallagma vernale, a new species of Odonata from Michigan. Occasional Papers of the Museum of Zoology, University of Michigan 479:1-8.

Hagen, H. A. 1861. Synopsis of the neuroptera of North America, with a list of the South American species. Smithsonia Miscellaneous Collections 4:1-347.

Ingham, B. R., and C. E. Jenner. 1976. Life histories of Enallagma hageni (Walsh) and E. aspersum (Hagen) (Zygoptera: Coenagrionidae). Odonatologica 5:331-345.

Johannsson, O. E. 1978. Co-existence of larval Zygoptera (Odonata) common to the Norfolk Broads (U.K.). Oecologia 32:303-321.

Kellicot, D. S. 1895. Catalogue of the Odonata of Ohio, Part 1. Journal of the Cincinnati Society of Natural History 17:195-216.

Kormondy, E. J., and J. L. Gower. 1965. Life history variations in an association of Odonata. Ecology 46:882-886.

Macan, T. T. 1964. The Odonata of a moorland fishpond. Int. Revue ges. Hydrobiol. 49:325-360.

Morse, A. P. 1895. New North American Odonata. Psyche 7:207-211.

O'Brien, M. F., and P. D. Pratt. 1999 (In press). Enallagma anna, a damselfly new to the Great Lakes region (Odonata: Coenagrionidae). The Great Lakes Entomologist 32(1).

Pearlstone, P. S. M. 1973. The food of damselfly larvae in Marion Lake, British Columbia. Syesis 6:33-39.

Say, T. 1839. Descriptions of new North American neuropterous insects and observations on some already described by (the late) Th. Say. Journal of the Academy of Natural Science of Philadelphia 8:9-46.

Selys-Longchamps, E. de. 1875. Notes on Odonata from Newfoundland collected in 1874 by Mr. John Milne. Entomologists Monthly Magazine 11:241-243.

Selys-Longchamps, E. de. 1876. Synopsis des agrionines, cinquième légion: Arion (suite). Le genre Agrion. Bulletin de l'Académie royale des Sciences de Belgique (2) 42:480-531.

Walker, E. M. 1953. The Odonata of Canada and Alaska, Vol. 1. University of Toronto Press: Toronto, Ontario. xi + 292 pp.

Walsh, B. D. 1862. List of the Pseudoneuroptera of Illinois contained in the cabinet of the writer, with descriptions of over forty new species, and notes on their structural affinities. Proceedings of the Entomological Society of Philadelphia 1862:361-402.

Walsh, B. D. 1863. Observations on certain N. A. neuroptera by Hagen, M. D., of Konigsberg, Prussia; translated from the original French MS., and published by permission of the author, with notes and descriptions of about twenty new N. A. species of Pseudoneuroptera. Proceedings of the Entomological Society of Philadelphia 2:167-272. [note: see also Walsh, B. D. 1862 for original description of adults, which were named in Walsh 1863. Source: Westfall and May 1996 ).

Westfall, M. J., Jr. and M. L. May. 1996. Damselflies of North America. Scientific Publishers: Gainesville, Florida. x + 650 pp.

Disclaimer: The Dragonflies of Sabah is an educational resource written largely by and for SMK (secondary school) students in Malaysia. TDS doesn't cover all species in Sabah, nor does it include all the latest scientific information. Though we edit our pages for accuracy, we cannot guarantee all information in these pages. While we provide references to books and websites that we believe are reputable, we cannot necessarily endorse the contents of references beyond our control.

NEW INDEX : Dragonfly     May 10, 2008 01:21:01 PM

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